Xizang (Tibet): Jiangda (Gyamda) Xian.
Narrow gorge ca. 6 km E in a straight line from the center of the city of Jiangda (Gyamda).
Logged Picea forest, now dominated by Betula. Slopes and meadows; slopes with numerous, scattered limestone boulders and diverse herbaceous flora in organically rich, dark soil and with deep moss layer in many places
Mostly under overhanging cliffs and boulders in moss and in soil.Fronds usually widely spaces, rarely a few clustered; rhizomes brittle.
D. E. Boufford, B. Bartholomew, D. A. Eaton, X. H. Li, R. H. Ree, B. Xu, J. P. Yue, J. W. Zhang & X. X. Zhu